• UNIT 2 CONCEPT OF ECOSYSTEM

    UNIT 2: CONCEPT OF ECOSYSTEM

    Key Unit Competence
    Describe the different components of an ecosystem, biogeochemical cycles and

    how energy flows in an ecosystem.

    Learning objectives

    By the end of this unit, I should be able to:

    – Describe an ecosystem
    – State the types and properties of an ecosystem
    – Describe the main components of an ecosystem
    – Explain the ecological factors influencing the life of organisms in an ecosystem
    – Define the terms: population, community, ecosystem, biome, niche and
        biosphere
    – Distinguish among; individuals, populations, communities, niche, habitat,
        ecosystems, biomes, biosphere
    – Describe feeding relationships in an ecosystem
    – Describe a food chain and a food web
    – Explain the relative merits of pyramids of numbers
    – Analyse the relation between organisms (example: producers, consumers,
        decomposers) and their trophic levels.
    – Distinguish between abiotic and biotic factors
    – Interpret energy flow diagrams
    – Compare; gross primary, net primary production and secondary succession in
        biotic communities
    – Explain what is meant by trophic efficiency
    – Explain energy flow and the recycling of nutrients in an ecosystem
    – Describe biogeochemical cycles
    – Identify processes, components, and roles of organisms in the hydrologic,
        carbon and nitrogen cycles
    – Distinguish between primary and secondary succession in biotic communities
    – Appreciate the existence of different components of an ecosystem and their
        roles in the life of organisms
    – Beware of the effect of bioaccumulations at different trophic levels.

    – Recognise the source and transfer of energy in an ecosystem

    Introductory activity
    The following pictures indicate different types of ecosystems. Observe

    carefully the pictures A, B and C and answer the questions that follow.

    1. What do you understand by the terms: ecosystem, biotic and abiotic factors?
    2. Suggest the types of ecosystems illustrated by pictures A, B, and C.
    3. Distinguish between abiotic and biotic factors illustrated on picture A, B and
        C.
    4. Describe how energy flows through ecosystem B and ecosystem C.
    5. Explain how feeding relationships are expressed in food chains on picture B
         and C.
    6. Identify trophic levels in food chains and food webs on the picture B and
        picture C.
    7. What would happen if plant species are removed from an ecosystem of

        picture C?

    Ecology is the study of how living things interact with each other and with their
    environment. It is one of the major branches of biology with different areas that
    overlap with geography, geology, climatology, mathematics, and chemistry cited
    among other sciences. This lesson introduces fundamental concepts in ecology
    with a particular focus on organisms and their environment. Organisms are
    individual living things. Despite their tremendous diversity, all organisms have
    the same basic needs such as energy and matter, obtained from the environment.
    Therefore, organisms are not closed systems. They depend on and are influenced
    by the environmental factors including abiotic (non-living factors such as water,
    temperature, humidity…) and biotic (living factors such as animals, plants…). The
    unit of nature consisting of all the biotic and abiotic factors in an area and their

    interactions is called an ecosystem.

    2.1 Ecosystem
    Activity 2.1

    Observe carefully the diagram below, and answer the questions that follow

    1. Define an ecosystem and give its different types.
    2. Distinguish among; individuals, populations, communities, niche,

         habitat, ecosystems, biomes and the biosphere.

    Different concepts define levels in ecology. From the low to high level, the concepts

    include:

    a. Species
    Species such as bees in figure 2.1 is defined as a group of organisms that can breed

    to produce fully fertile offspring.


                                                                                        Figure 2.1: Species of bees

    b. Population
    A population is defined as a group of organism of the same species which live in the
    same habitat at the same time where they can freely interbreed. Elephants such as

    those indicated in figure 2.2 constitute a population.


                                                                                         Figure 2.2: Population of elephants

    c. Community
    In ecology, a community consists of all populations of different species living and
    interacting at a certain level in the same ecosystem. Animals indicated in the figure

    2.3 interact and share the same ecosystem


                                                                        Figure 2.3: Ecological community

    d. Niche
    A niche refers to the role played by a species in its ecosystem. It includes all the ways
    that the species interacts with the biotic and abiotic factors of the environment.
    Two important aspects of a species’ niche are the food it eats and how the food is
    obtained. Birds on the figure 2.4 live in the same ecosystem, but they have different
    adaptations for food. For example, the longest slender beak of the nectarivore allows
    it to sip the nectar from flowers, the short study beak of the granivore allows it to

    crush hard and tough grains.


    Figure 2.4: Adaptations of birds’ beak for food in an ecosystem

    Another aspect of a species’ niche is its habitat. The habitat is the physical environment
    in which a species lives and to which it is adapted. A habitat’s features are mainly
    determined by abiotic factors such as temperature and rainfall, which in turn have
    an influence on the traits of the organisms that live in that habitat. A habitat is also
    influenced by biotic factors as it may contain many different species. However, in the
    same habitat, two different species cannot occupy the same niche in the same place
    for very long. This is known as the competitive exclusion principle. If two species
    were to occupy the same niche, they would compete with one another for the same
    food and other environmental resources leading to the extinction of the weaker

    species.

    e. Ecosystem
    An ecosystem consists of a natural unit consisting of all the living organisms in an
    area functioning together with all the non-living physical factors of the environment.
    The concept of an ecosystem can apply to units of different sizes. For example, a
    large body of fresh water could be considered an ecosystem, and so could a small
    piece of dead wood. Both contain a community of species that interact with one

    another and with the abiotic components of their environment.

                                                                        Figure 2.5: Example of ecosystems

    They are two major classification of ecosystems: natural ecosystem and artificial
    ecosystem. Natural ecosystems are those ecosystems that are capable of operating
    and maintaining themselves without any major interference by man. Natural
    ecosystems are furthermore classified into terrestrial ecosystems including; forest,
    grassland and desert, and in Aquatic ecosystems including fresh water ecosystem

    such as; ponds, lakes, rivers and marine ecosystems such as ocean, sea or estuary.

    Artificial Ecosystem are those ecosystems maintained by the intervention of humans.
    They are manipulated by man for different purposes including; croplands, artificial

    lakes and reservoirs, townships and cities.


                                                      Figure 2.6: Artificial ecosystem

    f. Biomes
    A biome is a broad regional type of an ecosystem characterized by distinctive climate
    and soil conditions and a distinctive kind of biological community adapted to those
    conditions. Biomes are of various types including terrestrial and aquatic biomes.
    Terrestrial biomes consist of all the land areas on Earth where organisms live. The
    distinguishing features of terrestrial biomes are determined mainly by climate.
    The dominant terrestrial biomes include; tundra, temperate forests, grasslands,

    temperate, tropical deserts, tropical forests and grasslands (Figure 2.7).

                                                                         Figure 2.7: Different types of biomes

    Aquatic biomes occupy the largest part of biosphere. These are divided into two,
    i.e. marine and freshwater. The marine biomes e.g. oceans which is the biggest
    of the two (Figure 2.8 below) have a very high salt concentration and have fauna
    adapted to them. The fresh water biomes such as lakes and rivers have a low salt

    concentration of less than 1%.


                                                                             Figure 2.8: An example of aquatic biome

    g. Biosphere
    The biosphere is the portion of Earth inhabited by life and which represents the sum

    of all communities and ecosystems.

    Application 2.1
    1. Distinguish among; individuals, populations, communities, ecosystems,
        biomes and biosphere.
    2. Give an example of any three aquatic and three terrestrial ecosystems
        found in Rwanda
    3. Use the examples above and make a brief description of an ecosystem

    4. Discuss the competitive exclusion principle.

    2.2 Properties of an ecosystem and ecological factors

           influencing the life of organisms

    Activity 2.2
    1. Go to your school garden and collect 3 living things and 3 non living
        things
    2. Discuss differences and similarities between collected living and nonliving
        things
    3. Analyze carefully the diagram below and answer the questions that
        follow:

    Make a classification of living things by the letters A, B, C, D, E, F and G based
    on the principle of being eaten by

    2.2.1 Relationships in an ecosystem
    In an ecosystem, living things have feeding relationships. In terms of sources of food,
    organisms are classified as; producers, consumers, or decomposers.
    – Producers are organisms that can manufacture their own food. They include;
       green algae , green plants and other autotrophs that are able to make their own
       food through photosynthesis or chemosynthesis
    – Consumers are organisms that obtain food from other organisms because they
       cannot make their own food. Based on their level of feeding, consumers are
       classified as primary consumers when they feed directly on plants. Primary
       consumers include herbivorous or omnivorous animals. Consumers are
       also classified as secondary consumers, when they feed directly on primary
       consumers. Secondary consumers include carnivorous animals. Tertiary
       consumers are consumers that feed directly on secondary consumers and are
       top carnivorous or omnivorous animals.

    – Decomposers are organisms that break down the tissues of dead organisms
       into simpler substances, for example bacteria and fungi that break down dead
       plants and animals into compounds of carbon and nitrogen. These compounds
       are released into the soil to be used by plants and animals for growth.
    In a food chain, producers such as plants produce their own energy without
    consuming other life forms. They gain their energy from conducting photosynthesis
    via sunlight. Consumers exist on the next level of the food chain and they are three
    main types of consumers namely herbivores, carnivores and omnivores. Consumers

    get the energy by feeding on plants or by eating other carnivores or herbivores.

    2.2.2 The ecological factors influencing the life of organisms in an
               ecosystem
    In an ecosystem, life is influenced by biotic and abiotic factors.

    a. Abiotic factors

    Light: Light plays an important role in the species composition and development
    of vegetation. Light is abundantly received on the surface of the earth from solar
    energy and it is used by primary producers to do photosynthesis. Light intensity
    shows special variations due to the factors like atmospheric water layer, particles
    dispersed in the air, etc. Furthermore, the vegetation of an area may also affect the
    light intensity. In deep shade under trees, or under water, light becomes limiting

    factor below which photosynthesis is not sufficient for effective growth.

    Temperature: Temperature is a measurement of the degree of heat. Like light,
    heat is a form of energy. The radiant energy received from the sun is converted into
    heat energy. Heat is measured in calories. The temperature at which physiological

    processes are at their maximum efficiency is called optimum temperature.

    The minimum, optimum and maximum temperatures are called cardinal
    temperatures. The cardinal temperature varies from species to species and in the
    same individual from part to part. The distributions of plants, animals are also

    influenced by temperature.

    Water: Water is an indispensable part of land contributing to soil productivity, and
    the well beings of organisms. All physiological processes take place in the medium
    of water. For example, cellular protoplasm is made up mostly of water contributing

    to the maintenance of cells and hence the entire living organism survives.

    Rainfall: The rainfall provides water to plants and animals, and determines the
    types of vegetation in a given region. For example, the evergreen forests are found
    in tropical regions. Changes in rainfall influence the vegetation types in different
    parts of the earth, and in turn, vegetation causes changes in the types of forests,
    animals and birds. The quantity of water that a soil holds or that infiltrates into the
    soil depends upon the properties of soil and type and density of vegetation covering
    it. In a bare area, the rain drops beat the compact surface of the soil and loosen the

    soil particles which are washed away.

    Wind: Air in motion is called wind. It modifies the water relation and light conditions
    of a particular region, and brings about a number of physical, anatomical and
    physiological changes of plants. Such changes are breakage and uprooting of
    plants, deformation, erosion and deposition of different organic particles. The wind
    accelerates transpiration, removes solid moisture and at high velocities causes soil
    erosion, which contributes to the removal of the surface soil, rich in organic matter

    and fine mineral particles.

    Humidity: Humidity is greatly influenced by intensity of solar radiation, temperature,
    altitude, wind, and water status of soil. Low temperature causes higher relative
    humidity by decreasing the capacity of air for moisture. Processes as transpiration,

    absorption of water are influenced by atmospheric humidity.

    Atmospheric Gases: Some principal gases like nitrogen, oxygen, carbon-dioxide,
    helium, hydrogen, methane, and ozone are found in atmosphere. In addition to
    these gases, there are water vapor. Industrial gases, dust, smoke particles, microorganisms
    are present in the atmosphere. These gases have different influences on

    the environment and hence on the living things.

    Biotic Factors
    The biotic factors constitute the living organisms of the environment and their
    direct or indirect interactions. The population occurring together in an area interacts
    with each other in several ways including predation, competition for mating and for
    different natural resources including; food, water and oxygen.
    b. Edaphic Factors
    Edaphic factors deal with different aspects of soil, such as the structure and
    composition of soil, its physical and chemical features. A galaxy of complex factor
    constitutes the soil. Soil is usually defined as any part of earth’s crust in which plants
    root. The soil is constituted as a result of long-term process of complex interaction
    leading to the production of a mineral matrix in close contact with interstitial
    organic matter both living and dead organisms. Soil is composed of; mineral matter,
    soil organic matter or humus, soil water and soil solutions, and biological systems

    including bacteria, fungi, algae, protozoans and arthropods.

    Application 2.2
    1. Discuss the ecological factors driving the biodiversity of Akagera National
         Park.
    2. Discuss the relationship between plant diversity and soil composition.

    2.3. Energy flow in an ecosystem

    Activity 2.3

    Observe carefully the diagram below and answer the questions that follow.

    1. Discuss how the energy flows in the above food chain of living things.
    2. Indicate which living organisms above are consumers, decomposers in
        the figure.
    3. Discuss the role played by organism represented by the letter C.

    4. What would happen if A is removed from the food chain?

    Energy enters in an ecosystem in the form of sunlight or chemical compounds. Some
    organisms including plants and green algae use sunlight energy to make their own
    food. Other organisms get energy through food by eating producers or consumers

    or by decomposing producers and consumers.

    2.3.1 Food chains and food webs
    Food chains and food webs are diagrams that represent feeding relationships. They
    show who eats who. In this way, they model how energy and matter move through
    ecosystems.
    a. Food chains
    A food chain represents a single pathway through which energy and matter flow
    through an ecosystem. Food chains are generally simpler than what really happens

    in nature. Most organisms consume and are consumed by more than one species.

                                      Figure 2.9: Illustration of a food chain (Source shutterstock.com)

    b. Food Webs
    A food web represents multiple pathways through which energy and matter flow
    through an ecosystem. It includes many intersecting food chains. It demonstrates

    that most organisms eat, and are eaten, by more than one species.

                                                                      Figure 2.10: Illustration of the Food Web

    c. Trophic levels
    The feeding positions in a food chain or web are called trophic levels. The different
    trophic levels are defined in the table below (Table 2.1). All food chains and food
    webs have at least two or three trophic levels, the maximum being of four trophic
    levels. Many consumers feed at more than one trophic levels. Humans, for example,
    are primary consumers when they eat plants, secondary consumers when they eat
    meat from primary consumers, and are tertiary consumers when they eat meat of
    secondary consumers.
    Table: 2.1. Description of producers, primary, secondary and tertiary trophic

    levels


    2.3.2 Ecological pyramids
    Ecological pyramid is a graphical representation in the form of a pyramid showing
    the feeding relationships of groups of organisms. It is often represented in a way

    that the producers are at the bottom level and then proceeds through the various

    trophic levels in which the highest is on top. There are 3 types of ecological
    pyramids: pyramid of numbers, pyramid of biomass and pyramid of energy.

    Pyramid of numbers

    Pyramid of numbers is a graph representing the total number of individuals present
    at each trophic level. This type of pyramid can have two different forms depending
    on the number of organisms: upright and inverted. In an upright pyramid of numbers,
    the number of organisms generally decreases from the bottom to top. This generally
    occurs in grassland and pond ecosystems where plants occupy the base of the
    pyramid. An inverted pyramid of numbers, on the other hand, is just the opposite
    of the upright one. It is usually observed in tree ecosystems with the trees as the

    producers and the insects as consumers.

                                 Figure 2.11: Figure 2.11: illustration of the upright pyramid of numbers 

    d. Pyramid of biomass
    Biomass is defined as the amount of biomass per unit area product of the living
    material present in an organism and the total number of organisms present in a
    specific trophic level. In less complicated terms, it refers to the food available for
    the succeeding trophic level. A pyramid of biomass is a depiction of the amount of
    food available and how much energy is being passed on at each trophic level. Most
    the biomass that animals consume is used to provide the energy, converted to new

    tissues, or just remain undigested.

    Most of the time, pyramids of biomass are in a true pyramidal shape with biomass
    in the lower trophic levels are greater than the trophic levels above them. Like the
    pyramid of numbers, the pyramid of biomass can either have two forms: upright and
    inverted. Usually, terrestrial ecosystems are characterized by an upright pyramid of
    biomass having larger base for primary producers with the smaller trophic levels for
    consumers located at the top (figure 2.17). Aquatic ecosystems are the complete
    opposite as they will assume the inverted structure of the pyramid. This is because
    the phytoplankton producers with generally smaller biomass are located at the base
    while the consumers having larger biomass are located at the top of the pyramid

    (figure 2.18)

       Figure 2.12: Illustration of upright pyramid of biomass(left) and the inverted pyramid of biomass(right).

    In other words, the phytoplankton has a short turnover time, which means they have
    a small standing crop compared to their production. The turnover time is calculated

    by the following formula:

    2.3.3 Pyramid of energy
    The pyramid of energy shows the overall energy in the ecosystem and how much
    energy is required by organisms as it flows up the higher trophic levels. This pyramid
    shows that energy is transferred from lower trophic levels with more amount of energy
    (producers) to higher ones (consumers) and converted in the biomass. Therefore, it
    can be concluded that organisms found at the highest trophic levels of shorter food
    chains bear greater amount of energy than the ones found in longer ones. Unlike
    the first two ecological pyramids, the pyramid of energy is always illustrated in an
    upright position, with the largest energy carriers at the base. The pyramid shows the

    total energy stored in organisms at each trophic level in an ecosystem.

    Starting with primary consumers, each trophic level in the food chain has only 10
    percent of the energy of the level below it (Figure 2.18). The energy available at a

    given trophic level is measured in Kilojoules per square metre per year (kJm-2Y-1).

                                                             Figure 2.13: Illustration of the Pyramid of energy

    2.3.4 Limitations of ecological pyramids
    While the three ecological pyramids are highly specific to the aspect of ecosystem
    they want to describe, all of them still tend to overlook important aspects. Some of
    these limitations are the following:
    – These types of pyramids only are applicable in simple food chains and not for
         the food webs and they also do not consider the possible presence of the same
         species at different trophic levels.
    – None of the three ecological pyramids provide any idea related to variations in
        seasons and climates.
    – Other organisms like microorganisms and fungi are not given specific role in

         the pyramids despite their vital roles in ecosystems.

    Application 2.3
    1. All scientists agree that the activities of living organisms play an important
        role in driving biogeochemical cycles, and that organism shape their
       environment to a considerable extent.
    a. Explain how, herbivores affect their grassland environment.
    b. What would happen if herbivores were removed from Akagera National
        Park?
    c. What would happen to Akagera National Park if overgrazing occurs?
    2. Explain why is only small portion of the solar energy that strikes Earth’s
        atmosphere stored by primary producers.
    3. The diagrams A, B, C and D indicate different cases of pyramid of numbers.

         Using your knowledge on pyramids, analyses and interpret each diagram

    4. Discuss the reasons why the transfer of energy in an ecosystem is referred to

        as energy flow, not as energy cycling.

    2.4 Ecological succession

    Activity 2.4
    In pair discuss the following:
    1. What happen to a but a month after bush fire?
    2. What would happen to your school basketball playground after 1, 5, 50,

         500 years if it was completely abandoned?

    Communities are not usually static, and the numbers and types of species that live in
    them generally change through time. This is called ecological succession. Important
    cases of succession are primary and secondary succession.
    a. Primary succession
    Primary succession occurs in an area that has never been colonized such as bare
    rock. This type of environment may come about when lava flows from a volcano and
    hardens into rock, a glacier retreats and leaves behind bare rock or when a landslide
    uncovers an area of bare rock.
    The first species to colonize a disturbed area are called pioneer species including
    bacteria and lichens that can live on bare rock. These species change the environment
    and make the way for other species to come into the area. Along with wind and
    water, they help weather the rock and form soil. Once soil begins to form, plants can
    move in from pioneer species to intermediate stages and to climax communities
    (Figure 2.14). At first, the plants include herbs, grasses and other species that can
    grow in thin, poor soil. As more plants grow and die, organic matter is added to the
    soil. Soil is improved and get the capacity to hold water. The improved soil allows

    shrubs and trees to move into the area.

                                                                               Figure 2.14: Primary succession

    b. Secondary succession
    Secondary succession occurs in a formerly inhabited area that was disturbed. The
    disturbance could be a fire, flood, or human action such as farming. This type of
    succession is faster because the soil is already in place. In this case, the pioneer
    species are plants such as grasses, birch trees, and fireweed. Organic matter from

    the pioneer species improves the soil and lets other plants move into the area.


                                                                          Figure 2.15: Secondary succession

    Similarities and differences between primary and secondary succession are
    summarized in the following table:

    Table: 2.2 Comparison between primary succession and secondary succession


    Application 2.4

    Differentiate between primary and secondary succession

    2.5 Bioaccumulation and Bio magnification
    Activity 2.5
    Use the school library and search additional information on the internet.

    Discuss between bioaccumulation and bio magnifications

    2.5.1 Bioaccumulation
    Bioaccumulation refers to the accumulation of toxic chemical substances such as
    pesticides, or other chemicals in the tissue of a particular organism. Bioaccumulation
    occurs when an organism absorbs a substance at a rate faster than that at which the

    substance is lost by catabolism and excretion

    2.5.2 Bio magnification
    Bio magnification is a process by which chemical substances become more
    concentrated at each trophic level. Bioaccumulors of toxic substances such as heavy
    metals and polychlorinated biphenyls that slowly increases up in concentration in
    living organisms including bacteria, algae, fungi, and plants.Bioaccumulants enter
    a body through contaminated air, water, and/or food, and keep on accumulating
    because they are very either slowly metabolized, not all metabolized, or are excreted

    very slowly

    2.5.3 Example of the causes of bio magnification
    Some toxic chemicals were deliberately put in the environment to kill insect pests.
    One of these pesticides is Dichloro Diphenyl Trichloroethane (DDT), which was
    used to control mosquitoes and other insect pests. It was commonly sprayed on
    plants and eventually entered water supplies. There it was absorbed by microscopic
    organisms, which in turn were eaten by small fish and the small fish eaten by larger
    fish from where it could have transferred to other animals, where it accumulates in
    the fat tissue of animals at the top of the food chain. This food chain shows typical

    concentrations of DDT found in a food chain (in parts per million, ppm):

    Another biological magnification of Polychlorinated Biphenols (PCBs) was found in
    the food web of great lakes, where the concentration of PCBs in herring gull eggs, at
    the top of the food web, is nearly 5,000 times that in phytoplankton at the base of

    the food web.

                                  Figure 2.16: Biological magnification of PCBs in a Great Lakes food web.

    2.5.4 Consequences of bio magnification
    The first sign of the problem was a decline in the number of predator birds. Studies
    showed that the eggs of these birds were easily cracked. In fact, the weight of the
    mother sitting on the eggs cracked them. It was finally discovered that DDT was
    building up in the tissue of the birds and interfering with the calcium needed for the

    shell to be hard.


                                         Figure 2.17: Biomagnification of pesticides in food chain

    2.5.5 Relationship between bioaccumulation and bio magnification


         Figure 2.18: Differences and similarities between bioaccumulation and bio magnification

    2.5.6 Prevention and reduction of bioaccumulation of toxic substances
    The following are some of the ways to prevent and to reduce bioaccumulation of
    toxic substances:
    – Do not put harmful substances into water system or storm drains.
    – Reduce the use of toxic chemical pesticides.
    – Eat certified organic foods when possible.

    – Avoid fishing or spending time in contaminated areas.

    Application 2.5
    1. Discuss how the addition of excess nutrients to a lake threatens the
        population of fishes.
    2. In the face of biological magnification of toxins such as DDT, discuss the

        levels of food chains where it is healthier to feed on

    2.6 Efficiency of ecological production
    Activity 2.6
    Use the books from the school library and search further information from
    the internet. Discuss the roles of efficiency of ecological production and
    make a brief description of the ecosystem primary production, total primary

    production, and net primary production.

    2.6.1 Efficiency of primary production
    The amount of light energy converted to chemical energy in the form of organic
    compounds by autotrophs during a given period of time is called ecosystem
    primary production (R). Most primary producers use light energy to synthesize
    energy rich-organic molecules, which are subsequently broken down to generate
    adenosine triphosphate (ATP). The total primary production in an ecosystem’s gross
    production (GPP) is the amount of light energy that is converted to chemical energy
    by photosynthesis per unit time.
    Note that not all of this production is stored as organic material in the primary
    producers because they use some of the molecules as fuel in their own cellular
    respiration. The net primary production (NPP) equals the gross primary production
    minus the energy used by the primary producers for respiration(R), as it is summarized

    in the following formula, i.e

    NPP = GPP – R.
    In many ecosystems, NPP is about one-half of GPP.
    To an ecologist, net primary production is the key measurement because it represents

    the storage of chemical energy that will be available to consumers in the ecosyste

                                                Figure 2.19: Illustration of the net primary productivity

    2.6.2 Efficiency of secondary production
    The amount of chemical energy in consumer’s food that is converted to their own
    biomass during a given period of time is called the secondary production of the
    ecosystem. Consider the transfer of organic matter from primary producers to
    herbivores, the primary consumers. In most ecosystems, herbivores eat only a small
    fraction materials produced by plants. Moreover, they cannot digest all the eaten
    plant materials. Thus, much of primary production is not used for consumers. In this

    case, the secondary production is calculated by:

    Net Secondary Production (NSP) = Gross Secondary Production (GSP) – Respiration(R)


                                                        Figure 2.20: Net secondary production

    2.6.3 Ecological production efficiency

    Production efficiency is the percentage of energy stored in assimilated food that is

    not used for respiration. It is calculated as follows:


    Production efficiency is expressed in percentage (%)
    As an example, when a caterpillar feeds on a plant leaf, only about 33 J of out 200 J, or one-sixth
    of the energy in the leaf is used for secondary production or growth. The caterpillar uses some of
    the remaining energy for cellular respiration and passes the rest in faeces. The energy contained in
    faeces remains in the ecosystem temporarily, but most of it is lost as heat after the faeces are
    consumed by detritivores. The energy used for caterpillar’s respiration is also lost from the

    ecosystem as heat.


    Application 2.6
    1. As part of a new reality show on television, a group of overweight people are
         trying to safely lose in one month as much weight as possible. In addition to
        eating less, what could they do to decrease their production efficiency for

        the food they eat?

    2. Tobacco leaves contain nicotine, a poisonous compound that is energetically
        expensive for the plant to make. What advantage might the plant gain by
        using some of its resources to produce nicotine?
    3. If an insect eats plant seeds containing 100J of energy, energy from which 30
         J is used for respiration while 50J remains in faeces.
    4. a. Calculate the net secondary production.

         b. Estimate the production efficiency.

    2.7 Biogeochemical Cycles
    Activity 2.7

    Observe carefully the diagrams below and answer the questions that follow.


    1. Name the biogeochemical cycles represented by X, Y and Z.
    2. For the biogeochemical cycles denoted X, Y and Z, make a description of
         steps represented by the letters A, B and C.
    3. What do you understand by the term biogeochemical cycle?

    4. Discuss the importance of biogeochemical cycles to living things e.g. man.

    A biogeochemical cycle is a closed loop through which a chemical element or
    water moves through ecosystems. In the term biogeochemical, bio- refers to
    biotic components and geo- to geological and other abiotic components. During
    biogeochemical cycle, chemicals cycle through both biotic and abiotic components
    of ecosystems. For example, an element might move from the atmosphere to the
    water of the ocean, goes to ocean organisms, and then back to the atmosphere to

    repeat the cycle.

    Elements or water may be held for various lengths of time by different components
    of a biogeochemical cycle. Components that hold elements or water for a relatively
    short period of time are called exchange pools. For example, the atmosphere is
    an exchange pool for water. It holds water for several days. This is a very short time
    compared with the thousands of years the deep ocean can hold water. The ocean
    is an example of a reservoir for water. A reservoir is a component of a geochemical

    cycle that hold elements or water for a relatively longer period of time.

    2.7.1 Water Cycle
    Earth’s water is constantly in motion. Although the water on Earth is billions of years
    old, individual water molecules are always moving through the water cycle. The
    water cycle describes the continuous movement of water molecules on above and
    below Earth’s surface. Like other biogeochemical cycles, there is no beginning or
    end to the water cycle. It just keeps repeating. During the cycle, water occurs in its
    three different states: gas (water vapour), liquid (water), and solid (ice). Processes
    involved in changes of state in the water cycle include; evaporation, sublimation,

    and transpiration.

                                                                 Figure 2.21: Illustration of the water cycle

    2.7.2 Carbon Cycle
    Carbon is essential to all life as it is the main constituent of living organisms. It serves
    as the backbone component for all organic polymers, including; carbohydrates,
    proteins, and lipids. Carbon compounds such as carbon dioxide (CO2) and methane

    (CH4) circulate in the atmosphere and influence global climates. Carbon circulates

    between living and non-living components of the ecosystem primarily through
    the processes of photosynthesis and respiration. Plants and other photosynthetic
    organisms obtain CO2 from their environment and use it to build biological
    materials. Plants, animals, and decomposers (bacteria and fungi) return CO2 to the
    atmosphere through respiration. CO2 trapped in rock or fossil fuels can be returned
    to the atmosphere via volcanic eruptions, or fossil fuel combustion. The movement
    of carbon through biotic components of the environment is known as the fast
    carbon cycle.

                                                                                    Figure 2.22: The carbon cycle

    2.7.3 Nitrogen Cycle
    The atmosphere is the largest reservoir of nitrogen on Earth. It consists of 78%
    nitrogen gas (N2). Similar to carbon, nitrogen is a necessary component of biological
    molecules. Atmospheric nitrogen (N2) is converted to ammonia (NH3) by nitrogenfixing
    bacteria in aquatic and soil environments. These organisms use nitrogen to
    synthesize the biological molecules they need to survive. Some nitrogen-fixing
    bacteria live in soil, others live in the root nodules of legumes such as; peas and

    beans. In aquatic ecosystems, some cyanobacteria are nitrogen fixing.


                              Figure 2.23: Illustration of the nitrogen cycle (Adapted from Pearson Education, 2003)

    2.7.5 The Greenhouse Effect
    The greenhouse effect is a natural process that warms the Earth’s surface. When the
    sun’s energy reaches the Earth’s atmosphere, some of it is reflected back to space
    and the rest is absorbed and re-radiated by greenhouse gases. Greenhouse gases
    include water vapor, carbon dioxide, methane, nitrous oxide, ozone and some
    artificial chemicals such as chlorofluorocarbons (CFCs). The absorbed energy warms
    the atmosphere and the surface of the Earth. This process maintains the Earth’s
    temperature at around 330C warmer than it would otherwise be, allowing life on
    Earth to exist. The problem we now face is that human activities particularly burning
    fossil fuels (coal, oil and natural gas), agriculture and land clearing are increasing the
    concentrations of greenhouse gases. This is the enhanced greenhouse effect, which

    is contributing to the global warming.

    Application 2.7

    The diagram below shows the carbon cycle.

    Identify processes labelled ①, ② and ③.
    b. Describe two ways by which carbon can be removed from the cycle for
         long period of time.
    c. Describe two activities of humans that are disrupting the natural

         carbon cycle.

    End of unit assessment 2
    Section A: Multiple choice questions
    Choose the letter that best answers the question or completes the statement
    1. All of life on Earth exists in a region known as
       a. Ecosystem
       b. Biome
       c. Biosphere
       d. Ecology
    2. Groups of different species that live together in a defined area make up
       a. Population
       b. Community
       c. Ecosystem
       d. Biosphere
    3. The series of steps in which a large fish eats a small fish that has eaten algae
         is a) Food web b) Food chain c) Pyramid of numbers d) Biomass pyramid
    4. The total mass of living tissue at each trophic level can be shown in
       a. Energy pyramid
       b. Pyramid of numbers
       c. Biomass pyramid
       d. Biogeochemical cycle
    5. An ecosystem is not considered to be self-sustaining if
       a. There is interaction between biotic and abiotic factors
    b. Some of its living organisms incorporate energy into organic compounds
       c. Cycling of materials occurs between organisms and their environment
       d. It lacks a constant supply of energy
    Section B: Questions with short answers
    6. What is the meaning of the term ecology?
    7. Name the different levels of organization within the biosphere, from smallest
        to largest
    8. How is sunlight important to most ecosystems?
    9. By what process do:
        a. Decomposers convert organic matter into ammonia
        b. Bacteria convert gaseous nitrogen into ammonia
        c. Nitrosomonas convert ammonia into nitrites

        d. Pseudomonas convert nitrates into gaseous nitrogen

    10. Why is the transfer of energy and matter in a food chain only about 10
           percent efficient?
    Section C: Essay questions
    11. Describe the three different types of ecological pyramids.
    12. Why do the rectangles in a pyramid of energy get smaller at each higher
           trophic level?
    13. Discuss the reasons why the secondary succession is usually much faster
           than primary succession?
    14. The diagram below shows part of the nitrogen cycle
             a. Name a genus of bacteria which is responsible for each of the reactions
             A, B, C and D.
             b. Describe the conditions in which the bacteria responsible for reaction
             D will thrive.
    15. The table below shows mean values for primary productivity for four
           ecosystems: temperate deciduous forest, tropical forest, temperate grassland,

           and intensively cultivated land in a temperate region

    a. Suggest two reasons to account for the higher primary productivity of
         a tropical forest compared with a temperate forest.
    b. Suggest explanations for the difference in primary productivity
         between temperate grassland and intensively cultivated land.
    c. Describe how you would estimate the fresh biomass of the producers

          in a grassland ecosystem.

    16. The diagram shows a number of stages in an ecological succession in a lake.

    a. Use information from this diagram above and explain what is meant by
        an ecological succession.
    b. Give two general features this succession has in common with other
        ecological successions.
    c. A number of small rivers normally flow into the lake. These rivers flow
        through forested areas. Explain how deforestation may affect the process
        of succession in the lake.
    17. Use the skills learnt in classroom and give answers to the following questions:
    a. What is an ecosystem?
    b. What is the required information to fully describe the make-up of an
         ecosystem?
    c. Discuss the flow of energy through ecosystems and make a description of
         the various ways in which human activity can influence the energy flow at
         all levels in terrestrial ecosystems
    18. As part of a science project, Abingondo Diane is trying to estimate total primary
            production of plants in a prairie ecosystem for a period of one year. Once per
            quarter, Abingondo cuts a plot of grass with a lawnmower, do a collection and
            weighs the cuttings with the main purpose to estimate plant production. What

            is missing for Abingondo to estimate the total primary production?

    UNIT 1 POPULATION AND NATURAL RESOURCESUNIT 3 EFFECT OF HUMAN ACTIVITIES ON ECOSYSTEM